Quasi-steady aerodynamics predicts the dynamics of flapping locomotion

The propulsion of a flapping wing or foil is emblematic of bird flight and fish swimming. Previous studies have identified hallmarks of the propulsive dynamics that have been attributed to unsteady effects such as the formation and shedding of edge vortices and wing–vortex interactions. Here, we show that several key features of heaving flight are captured by a quasi-steady aerodynamic model that aims to predict stroke-averaged forces from wing motions without explicitly solving for the flows. We address the forward dynamics induced by up-and-down heaving motions of a thin plate with a nonlinear model which involves lift and drag forces that vary with speed and attack angle. Simulations reproduce the well-known transition for increasing Reynolds number from a stationary state to a propulsive state, where the latter is characterised by a Strouhal number that is conserved across broad ranges of parameters. Parametric, sensitivity and stability analyses provide physical interpretations for these results and show the importance of accounting for the flow regimes which are demarcated by Reynolds number and angle of attack. These findings extend the phenomena of unsteady locomotion that can be explained by quasi-steady modelling, and they broaden the conditions and parameter ranges over which such models are applicable.